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context:
Lenina chats in the bathroom with Fanny Crowne about her relationship with Henry Foster. Fanny chides Lenina for going out with Henry almost exclusively for four months, and Lenina admits she is attracted to the strange, somewhat funny-looking Bernard Marx. In another part of the Hatchery, Bernard is enraged when he overhears a conversation between Henry and the Assistant Predestinator about “having” Lenina.

further explanation:
There is a hatchery which produces human embryos and they are destined according to the will of controllers of the factory. One of the persons who helps the controllers is "Assistant Predestinator". In the story, he is called in this way.